Module 1: CELL STRUCTURE AND FUNCTION

Lecture 5: Structure and Function of Cytoplasm, Nucleus and Mitochondria

Properties of cytoplasmic matrix

The most of the physical properties of the matrix are due to its colloidal nature. The cytosol shows Tyndal effect (light scattering by particle in colloidal solution) and Brownian motion (random moving of particles). Due to the phase reversal property of the cytoplasmic matrix, the intracellular streaming or movement of the matrix takes place and is known as the cyclosis. The cyclosis usually occurs in the sol-phase of the matrix and is effected by the hydrostatic pressure, temperature, pH, viscosity, etc. Cyclosis has been observed in most animal and plant cells. The amoeboid movement depends directly on the cyclosis. The amoeboid movement occurs in the protozoans, leucocytes, epithelia, mesenchymal and other cells. Due to cyclosis matrix moves these pseudopodia and this causes forward motion of the cell. The cytoplasmic matrix being a liquid possesses the property of surface tension. The proteins and lipids of matrix have less surface tension, therefore, occur at the surface and form the membrane, while the chemical substances such as NaCl have high surface tension, therefore, occur in deeper part of the matrix. Besides surface tension and adsorption, the matrix possesses other mechanical properties, e.g., elasticity, contractility, rigidity and viscosity which provide to the matrix many physiological utilities. The colloidal system due to its stable phase gives polarity of the cell matrix which cannot be altered by centrifugation of other mechanical means. The matrix has a definite pH value and it does not tolerate significant variations in its pH. Yet various metabolic activities produce small amount of excess acids or bases which is maintained by certain chemical compounds as carbonate-bicarbonate buffers. The matrix is a living substance and possesses various biological properties as irritability, conductivity, movement, metabolic activity, growth and reproduction.

Organelles
Cytoplasm contains all the organelles like nucleus, mitochondria, endoplasmic reticulum, lysosomes and Golgi apparatus. Besides, it also contains chloroplast in plant cells. Each organelle is bounded by a lipid membrane, and has specific functions.

Cytoplasmic inclusions
Some insoluble suspended substances found in cytosol. They are basically granules of starch and glycogen, and they can store energy. Besides, crystals of some minerals and lipid droplets can also be found in cytoplasm. Lipid droplets act as storage site of fatty acid and steroids.

Functions of Cytoplasm

Cytoplasm is the site of many vital biochemical reactions crucial for maintaining life.

  1. It is the place where cell expansion and growth take place.
  2. It provides a medium in which the organelles can remain suspended.
  3. Besides, cytoskeleton found in cytoplasm gives the shape to the cell, and facilitates its movement.
  4. It also assists the movement of different elements found within the cell.
    The enzymes found in the cytoplasm breaks down the macromolecules into small parts so that it can be easily used by the other organelles like mitochondria. For example, mitochondria cannot use glucose present in the cell, unless it is broken down by the enzymes into pyruvate. They act as catalysts in glycolysis, as well as in the synthesis of fatty acid, sugar and amino acid.
  5. Cell reproduction, protein synthesis, anaerobic glycolysis, cytokinesis are some other vital functions that are carried out in cytoplasm.

    However, the smooth operation of all these functions depend on the existence of cytoplasm, as it provides the medium for carrying out these vital processes.